A woman buried in Germany in the Late Neolithic era

The Late Neolithic period in Alberstedt, Germany, is part of the broader European Neolithic era, which marked a time of significant transformation in human societies across Europe. This period, which occurred approximately between 3000 and 2000 BCE, was characterized by the development of agriculture, advancements in tool-making, and early societal organization. Let’s explore the different aspects that define the Alberstedt region during this crucial period.
Geographic and Environmental Context
Alberstedt is located in the region of modern-day Saxony-Anhalt, Germany. This area is characterized by fertile loess soils, which provided an ideal environment for Neolithic farmers to practice agriculture. The region’s temperate climate with distinct seasons allowed for the cultivation of a variety of crops and supplemented by natural resources such as forests, rivers, and wildlife, offering a balanced diet and materials for tools and construction.
Subsistence and Economy
The economy of Late Neolithic Alberstedt was primarily agrarian. Archaeological evidence indicates that the inhabitants practiced mixed farming, which included the cultivation of cereals such as wheat and barley, and the domestication of animals like cattle, sheep, and pigs. The agricultural surplus eventually led to the development of trade networks with neighboring communities, which facilitated the exchange of goods, ideas, and technologies.
In addition to agriculture, the community relied on hunting, fishing, and gathering to supplement their diet. Access to nearby water bodies provided fish and other aquatic resources, while forests offered game and gatherable plants and nuts.
Settlements and Architecture
Settlements during the Late Neolithic in Alberstedt consisted of small, semi-permanent villages. The architecture predominantly comprised longhouses built using timber and wattle-and-daub techniques. These structures were not only homes but also served as communal spaces for activities like storage, craft production, and social gatherings.
Villages were typically strategically located near water sources and fertile land to ensure easy access to resources. As populations grew, these settlements became more complex, possibly leading to the formation of proto-urban centers.
Social Structure and Culture
Social organization in Late Neolithic societies was starting to become more complex, with evidence pointing towards emerging hierarchies. The division of labor likely existed, with specific roles such as farmers, hunters, toolmakers, and possibly religious or spiritual leaders.
Cultural practices included ceremonies and rituals, which were often linked to the agricultural calendar. Artefacts such as pottery, tools, and figurines suggest a developed material culture. Pottery styles, in particular, provide insight into the cultural and trade connections with other Neolithic groups across Europe.
Technology and Craft
Technological advancements during this period were significant. The people of Alberstedt were skilled in flintknapping, producing a range of tools for farming, hunting, and domestic purposes. The development of polished stone tools, such as axes and sickles, facilitated more efficient agricultural and forestry activities.
In pottery, the Late Neolithic period saw the emergence of more sophisticated techniques and designs. Pottery from Alberstedt featured both functional and decorative elements, indicative of the cultural identity and artistic expression of the people.
Spiritual and Religious Beliefs
While specific beliefs are hard to reconstruct fully, the presence of burial sites and ritual objects suggests that the Alberstedt community had a rich spiritual life. Grave goods and the orientation of graves indicate beliefs in an afterlife or ancestral veneration. Megalithic structures found in the region imply communal and possibly religious activities, reflecting a complex belief system tied to natural cycles and possibly celestial observations.
Conclusion
The Late Neolithic period in Alberstedt, Germany, represents a dynamic and transformative era. It is marked by the beginnings of organized agriculture, settlement, and societal structures, laying the groundwork for future developments in European prehistory. Archaeological findings from this region provide valuable insights into the daily lives, social fabric, and cultural practices of its inhabitants, offering a glimpse into the evolutionary path from nomadic to semi-permanent settlement lifestyles.
Sample ID | Culture/Period | Date | Location | Action |
---|---|---|---|---|
I0234 | Srubnaya Culture | 1850 BCE | Samara Oblast. Volga River Valley. Samara Steppes. Rozhdestveno I, Russia | View |
I0126 | Middle Bronze Poltavka | 2867 BCE | Samara Oblast. Volga River Valley. Kutuluk River. Kutuluk III, Russia | View |
I0432 | Middle Bronze Poltavka | 2925 BCE | Samara Oblast. Sok River. Potapovka I, Russia | View |
I0434 | Eneolithic Khvalynsk Culture, Russia | 5198 BCE | Samara Oblast. Volga River Valley. Khvalynsk II, Russia | View |
I0433 | Eneolithic Khvalynsk Culture, Russia | 4697 BCE | Samara Oblast. Volga River Valley. Khvalynsk II, Russia | View |
I0122 | Eneolithic Khvalynsk Culture, Russia | 4936 BCE | Samara Oblast. Volga River Valley. Khvalynsk II, Russia | View |
I0124 | Hunter-Gatherer Samara, Russia | 5660 BCE | Samara Oblast. Volga River Valley. Sok River. Lebyazhinka, Russia | View |
I0370 | Early Bronze Samara Yamnaya | 3300 BCE | Samara Oblast. Volga River Valley. Eastern Orenburg. Pre-Ural steppe. Ishkinovka I, Russia | View |
I0441 | Early Bronze Samara Yamnaya | 3010 BCE | Samara Oblast. Volga River Valley. Buzuluk. Kurmanaevka III, Russia | View |
I0444 | Early Bronze Samara Yamnaya | 3335 BCE | Samara Oblast. Volga River Valley. Kutuluk River. Kutuluk, Russia | View |
I0439 | Early Bronze Samara Yamnaya | 3322 BCE | Samara Oblast. Volga River Valley. Sok River. Lopatino I, Russia | View |
I0357 | Early Bronze Samara Yamnaya | 3093 BCE | Samara Oblast. Volga River Valley. Sok River. Lopatino I, Russia | View |
I0429 | Early Bronze Samara Yamnaya | 3339 BCE | Samara Oblast. Volga River Valley. Sok River. Lopatino I, Russia | View |
I1282 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1302 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1276 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1284 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1274 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1280 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1314 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1277 | Chalcolithic Spain | 2570 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1272 | Chalcolithic Spain | 2857 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1281 | Chalcolithic Spain | 2867 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1300 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I0807 | Middle Neolithic Baalberge Culture, Germany | 3977 BCE | Esperstedt, Germany | View |
I0559 | Middle Neolithic Baalberge Culture, Germany | 3646 BCE | Quedlinburg Site IX, Germany | View |
I0560 | Middle Neolithic Baalberge Culture, Germany | 3637 BCE | Quedlinburg Site IX, Germany | View |
I1546 | Bell Beaker Culture, Germany | 2500 BCE | Benzingerode-Heimburg, Germany | View |
I0806 | Bell Beaker Culture, Germany | 2433 BCE | Quedlinburg Site VII, Germany | View |
I0805 | Bell Beaker Culture, Germany | 2465 BCE | Quedlinburg Site VII, Germany | View |
I0113 | Bell Beaker Culture, Germany | 2396 BCE | Quedlinburg Site XII, Germany | View |
I0112 | Bell Beaker Culture, Germany | 2455 BCE | Quedlinburg Site XII, Germany | View |
I1530 | Bell Beaker Culture, Germany | 2456 BCE | Rothenschirmbach, Germany | View |
I0111 | Bell Beaker Culture, Germany | 2474 BCE | Rothenschirmbach, Germany | View |
I0108 | Bell Beaker Culture, Germany | 2575 BCE | Rothenschirmbach, Germany | View |
I0171 | Late Neolithic Benzigerode-Heimburg, Germany | 2288 BCE | Benzingerode-Heimburg, Germany | View |
I0059 | Late Neolithic Benzigerode-Heimburg, Germany | 2343 BCE | Benzingerode-Heimburg, Germany | View |
I1542 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I1536 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I1544 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I1538 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I1539 | Corded Ware Culture, Germany | 2630 BCE | Esperstedt, Germany | View |
I0106 | Corded Ware Culture, Germany | 2461 BCE | Esperstedt, Germany | View |
I1540 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I1541 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I1532 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I0049 | Corded Ware Culture, Germany | 2463 BCE | Esperstedt, Germany | View |
I0211 | Hunter-Gatherer Karelia, Russia | 7050 BCE | Karelia. Yuzhnyy Oleni Ostrov, Russia | View |
I0061 | Hunter-Gatherer Karelia, Russia | 7050 BCE | Karelia. Yuzhnyy Oleni Ostrov, Russia | View |
I0550 | Late Neolithic Karsdorf, Germany | 2572 BCE | Karsdorf, Germany | View |
I0797 | Early Neolithic Linear Pottery Culture, Germany | 5500 BCE | Karsdorf, Germany | View |
I0795 | Early Neolithic Linear Pottery Culture, Germany | 5215 BCE | Karsdorf, Germany | View |
I0176 | Middle Neolithic Linear Pottery Culture, Hungary | 5209 BCE | Szemely-Hegyes, Hungary | View |
I0551 | Middle Neolithic Salzmuende Culture, Germany | 3400 BCE | Salzmünde-Schiepzig, Germany | View |
I0409 | Early Neolithic Spain | 5312 BCE | Els Trocs, Spain | View |
I0411 | Early Neolithic Spain | 5298 BCE | Els Trocs, Spain | View |
I0405 | Middle to Late Neolithic Spain | 3900 BCE | La Mina, Spain | View |
I0174 | Early Neolithic Starčevo Culture 1, Hungary | 5712 BCE | Alsonyek-Bataszek. Mérnöki telep, Hungary | View |
I0115 | Early Bronze Age Unetice Culture, Germany | 1959 BCE | Esperstedt, Germany | View |
I0117 | Early Bronze Age Unetice Culture, Germany | 2276 BCE | Esperstedt, Germany | View |
I0804 | Early Bronze Age Unetice Culture, Germany | 2139 BCE | Eulau, Germany | View |
I0803 | Early Bronze Age Unetice Culture, Germany | 2135 BCE | Eulau, Germany | View |
I0164 | Early Bronze Age Unetice Culture, Germany | 2027 BCE | Quedlinburg Site VIII, Germany | View |
I0114 | Early Bronze Age Unetice Culture, Germany | 2141 BCE | Esperstedt, Germany | View |
I0022 | Early Neolithic Linear Pottery Culture, Germany | 5500 BCE | Baden-Württemberg. Stuttgart-Mühlhausen. Viesenhaeuser Hof, Germany | View |
I0026 | Early Neolithic Linear Pottery Culture, Germany | 5500 BCE | Baden-Württemberg. Stuttgart-Mühlhausen. Viesenhaeuser Hof, Germany | View |
I0013 | Hunter-Gatherer Motala, Sweden | 5966 BCE | Motala. Kanaljorden, Sweden | View |
I0011 | Hunter-Gatherer Motala, Sweden | 5721 BCE | Motala. Kanaljorden, Sweden | View |
I0015 | Hunter-Gatherer Motala, Sweden | 5967 BCE | Motala. Kanaljorden, Sweden | View |
I0012 | Hunter-Gatherer Motala, Sweden | 5715 BCE | Motala. Kanaljorden, Sweden | View |
I0014 | Hunter-Gatherer Motala, Sweden | 5885 BCE | Motala. Kanaljorden, Sweden | View |
I0017 | Hunter-Gatherer Motala, Sweden | 5722 BCE | Motala. Kanaljorden, Sweden | View |
I1508 | Early Neolithic Körös Culture, Hungary | 5716 BCE | Berettyóújfalu-Morotva-Liget, Hungary | View |
I1500 | Middle Neolithic Alföld Linear Pottery Culture, Hungary | 5301 BCE | Kompolt-Kigyoser, Hungary | View |
I1100 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1102 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1099 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1103 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1101 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1097 | Neolithic Turkey | 6420 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0744 | Neolithic Turkey | 6400 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1096 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1098 | Neolithic Turkey | 6419 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0708 | Neolithic Turkey | 6224 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0745 | Neolithic Turkey | 6387 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0746 | Neolithic Turkey | 6070 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0707 | Neolithic Turkey | 6225 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0709 | Neolithic Turkey | 6223 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0725 | Neolithic Turkey | 6400 BCE | Menteşe, Turkey | View |
I0736 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1499 | Middle Neolithic Alföld Linear Pottery Culture Bükk Group, Hungary | 5286 BCE | Garadna, Hungary | View |
I1502 | Early Bronze Age Makó Culture, Hungary | 2195 BCE | Kompolt-Kigyoser, Hungary | View |
I1497 | Late Chalcolithic Baden Culture, Hungary | 3320 BCE | Apc-Berekalya I, Hungary | View |
I1495 | Late Neolithic Lengyel Culture, Hungary | 4496 BCE | Apc-Berekalya I, Hungary | View |
I1498 | Middle Neolithic Alföld Linear Pottery Culture, Hungary | 5297 BCE | Debrecen Tocopart Erdoalja, Hungary | View |
I1507 | Early Neolithic Hunter-Gatherer Körös Culture, Hungary | 5788 BCE | Tiszaszolos-Domaháza, Hungary | View |
I1496 | Middle Neolithic Linear Pottery Culture, Hungary | 5211 BCE | Apc-Berekalya I, Hungary | View |
I1505 | Middle Neolithic Alföld Linear Pottery Culture, Hungary | 5210 BCE | Polgár-Ferenci hát. M3-31, Hungary | View |
I1271 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1303 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1549 | Bell Beaker Culture, Germany | 2500 BCE | Benzingerode-Heimburg, Germany | View |
I0407 | Middle to Late Neolithic Spain | 3900 BCE | La Mina, Spain | View |
I0025 | Early Neolithic Linear Pottery Culture, Germany | 5500 BCE | Baden-Württemberg. Stuttgart-Mühlhausen. Viesenhaeuser Hof, Germany | View |
I0247 | Questionable Iron Age Scythian, Russia | 385 BCE | Samara Oblast. Volga River Valley. Volga Steppes. Nadezhdinka, Russia | View |
I0423 | Srubnaya Culture | 1850 BCE | Samara Oblast. Volga River Valley. Samara River. Barinovka I, Russia | View |
I0443 | Early Bronze Samara Yamnaya | 3300 BCE | Samara Oblast. Volga River Valley. Sok River. Lopatino II, Russia | View |
I0726 | Neolithic Turkey | 6400 BCE | Menteşe, Turkey | View |
I0103 | Corded Ware Culture, Germany | 2617 BCE | Esperstedt, Germany | View |
I0172 | Middle Neolithic Esperstedt, Germany | 3363 BCE | Esperstedt, Germany | View |
I0408 | Middle to Late Neolithic Spain | 3895 BCE | La Mina, Spain | View |
I0412 | Early Neolithic Spain | 5309 BCE | Els Trocs, Spain | View |
I0054 | Early Neolithic Linear Pottery Culture, Germany | 5216 BCE | Unterwiederstedt, Germany | View |
I0104 | Corded Ware Culture, Germany | 2563 BCE | Esperstedt, Germany | View |
I0116 | Early Bronze Age Unetice Culture, Germany | 2137 BCE | Esperstedt, Germany | View |
I0118 | Late Neolithic Alberstedt, Germany | 2469 BCE | Alberstedt, Germany | View |
I0406 | Middle to Late Neolithic Spain | 3900 BCE | La Mina, Spain | View |
I0410 | Early Neolithic Spain | 5298 BCE | Els Trocs, Spain | View |
I0413 | Early Neolithic Spain | 5304 BCE | Els Trocs, Spain | View |
I0235 | Srubnaya Culture | 1850 BCE | Samara Oblast. Volga River Valley. Samara Steppes. Rozhdestveno I, Russia | View |
I0374 | Middle Bronze Poltavka | 2800 BCE | Samara Oblast. Volga River Valley. Samara River. Nikolaevka III, Russia | View |
I1583 | Neolithic Turkey | 6424 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0581 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I0440 | Middle Bronze Poltavka | 2887 BCE | Samara Oblast. Volga River Valley. Sok River. Lopatino II, Russia | View |
I0371 | Middle Bronze Poltavka | 2871 BCE | Samara Oblast. Volga River Valley. Sok River. Grachevka, Russia | View |
I0438 | Early Bronze Samara Yamnaya | 3020 BCE | Samara Oblast. Volga River Valley. Samara River. Luzkhi I, Russia | View |
I0418 | Middle Bronze Poltavka | 2131 BCE | Samara Oblast. Volga River Valley. Samara River. Utyevka VI, Russia | View |
I0099 | Late Bronze Age Halberstadt, Germany | 1202 BCE | Halberstadt-Sonntagsfeld, Germany | View |
I0047 | Early Bronze Age Unetice Culture, Germany | 2116 BCE | Halberstadt-Sonntagsfeld, Germany | View |
I1534 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I0060 | Bell Beaker Culture, Germany | 2404 BCE | Rothenschirmbach, Germany | View |
I0723 | Neolithic Turkey | 6008 BCE | Menteşe, Turkey | View |
I0724 | Neolithic Turkey | 6400 BCE | Menteşe, Turkey | View |
I0727 | Neolithic Turkey | 6400 BCE | Menteşe, Turkey | View |
I1580 | Neolithic Turkey | 6381 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1581 | Neolithic Turkey | 6386 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1585 | Neolithic Turkey | 6217 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1579 | Neolithic Turkey | 6221 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0854 | Neolithic Turkey | 6228 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0581 | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View | |
I0440 | 2887 BCE | Samara Oblast. Volga River Valley. Sok River. Lopatino II, Russia | View | |
I0418 | 2131 BCE | Samara Oblast. Volga River Valley. Samara River. Utyevka VI, Russia | View | |
I0438 | 3020 BCE | Samara Oblast. Volga River Valley. Samara River. Luzkhi I, Russia | View | |
I0585 | 5982 BCE | Leon. La Brana-Arintero, Spain | View | |
I1534 | 2500 BCE | Esperstedt, Germany | View | |
I1504 | 987 BCE | Ludas-Varjú dűlő, Hungary | View | |
I0099 | 1202 BCE | Halberstadt-Sonntagsfeld, Germany | View | |
I0047 | 2116 BCE | Halberstadt-Sonntagsfeld, Germany | View | |
I1579 | 6221 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View | |
I1581 | 6386 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View | |
I1580 | 6381 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View | |
I1585 | 6217 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View | |
I1583 | 6424 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View | |
I0727 | 6400 BCE | Menteşe, Turkey | View | |
I0724 | 6400 BCE | Menteşe, Turkey | View | |
I0854 | 6228 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View | |
I0723 | 6008 BCE | Menteşe, Turkey | View | |
I0371 | 2871 BCE | Samara Oblast. Volga River Valley. Sok River. Grachevka, Russia | View | |
I0060 | 2404 BCE | Rothenschirmbach, Germany | View |
Sample ID | Culture/Period | Date | Location | Action |
---|---|---|---|---|
I0118 | Late Neolithic Alberstedt, Germany | 2469 BCE | Alberstedt, Germany | View |
Ancient genetic admixture analysis compares the DNA profile of this individual (I0118) with present-day reference populations. These results show what percentage of the individual's genetic makeup resembles ancient populations from different geographic regions.
Modern genetic admixture analysis compares the DNA profile of this individual (I0118) with present-day reference populations. These results show what percentage of the individual's genetic makeup resembles modern populations from different geographic regions.
These results complement the ancient ancestry components shown in the previous section, offering a different perspective on the individual's genetic profile by comparing it with modern reference populations rather than prehistoric ancestral groups.
The G25 coordinates for the sample I0118 are as follows. You can analyze its admixture using G25 Studio.
I0118,0.12471942,0.12391042,0.05404184,0.0446995,0.0326892,0.01704002,0.00568034,0.00260242,0.00172192,-0.00335882,-0.00456382,0.00308026,-0.00861328,-0.00583004,0.01667162,-0.00457076,-0.0123797,0.00105732,0.0021595,0.00266894,0.00209106,0.00096428,-0.00157658,0.00978828,-0.00249352
Genome-wide patterns of selection in 230 ancient Eurasians
Ancient DNA makes it possible to observe natural selection directly by analysing samples from populations before, during and after adaptation events. Here we report a genome-wide scan for selection using ancient DNA, capitalizing on the largest ancient DNA data set yet assembled: 230 West Eurasians who lived between 6500 and 300 bc, including 163 with newly reported data. The new samples include, to our knowledge, the first genome-wide ancient DNA from Anatolian Neolithic farmers, whose genetic material we obtained by extracting from petrous bones, and who we show were members of the population that was the source of Europe's first farmers. We also report a transect of the steppe region in Samara between 5600 and 300 bc, which allows us to identify admixture into the steppe from at least two external sources. We detect selection at loci associated with diet, pigmentation and immunity, and two independent episodes of selection on height.